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NOUBHANI, Abdelmajid & Henri CAPPETTA:Les Orectolobiformes, Carcharhiniformes et Myliobatiformes (Elasmobranchii, Neoselachii) des Bassins a Phosphate du Maroc (Maastrichtien-Lutetien basal) -
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The Orectolobiformes, Carcharhiniformes and Myliobatiformes (Elasmobranchii, Neoselachii) of phosphate basins of Morocco (Maastrichtian-Early Lutetian. Systematics, biostratigraphy, evolution and faunal dynamics.The phosphate series of Morocco (Early Maastrichtian-Early Ypresian) is well known for its richness in selachian teeth studied by ARAMBOURG in an important monograph in 1952.
Newly collected material is particularly rich in new taxa which justify a reexamination of these faunas. The object of the present account is the revision of two shark orders (Orectolobiformes and Carcharhiniformes) and of one ray order (Myliobatiformes) which appearvery diversified and rich in new taxa.
13 new genera, one new subgenus and 52 new species are described and figured.
Orectolobiformes: Chiloscyllium meraense nov. sp., C. salvani nov. sp., Delpitoscyllium africanum nov. gen., Ganntouria variabilis nov. gen. nov. sp., Ginglymostoma botmaense nov. sp., G. chenanei nov. sp., G. erramii nov. sp., G. khouribgaense nov. sp., G. maroccanum nov. sp., G. pectinatum nov. sp., Hologinglymostoma jaegeri nov. gen. nov. sp., Metlaouia delpiti nov. gen. nov. sp., Palaeorhincodon daouii nov. sp., Plicatoscyllium gharbii nov. sp., P. youssoufiaense nov. sp., Pseudoginglymostoma erguitaense nov. sp., P. idiri nov. sp.
Carcharhiniformes: Casieria maghrebiana nov. gen. nov. sp., Danogaleus gueriri nov. gen. nov. sp., Foumtizia abdouni nov. gen. nov. sp., F. arba nov. gen. nov. sp., F. gadaensis nov. gen. nov. sp., Galeorhinus mesetaensis nov. sp., Khouribgaleus gomphorhiza nov. gen., Microscyliorhinus simplex nov. sp., Palaeogaleus dahmanii nov. sp., P. larachei nov. sp., P. sublaevis nov. sp., Porodermoides spanios nov. gen. nov. sp., Premontreia (Oxyscyllium) peypouqueti nov. subgen. nov. sp., Pteroscyllium lamranii nov. sp., Scyliorhinus entomodon nov. sp., S. ptychtus nov. sp., S. sulcidens nov. sp., Stenoscyllium priemi nov. gen. nov. sp., Triakis antunesi nov. sp., T. tanoutensis nov. sp.
Myliobatiformes: Coupatezia ambroggii nov. sp., C. boujoi nov. sp., C. chariri nov. sp., C. danica nov. sp., C. elevata nov. sp., C. laevis nov. sp., C. larivei nov. sp., C. melittiana nov. sp., C. reniformis nov. sp., Dasyatis datasi nov. sp., D. martini nov. sp., D. ponsi nov. sp., Delpitia reticulata nov. gen. nov. sp., Heterobatis talbaouii nov. gen. nov. sp., Heterotorpedo brahimi nov. sp., Heterotorpedo lyazidii nov. sp., Myliodasyatis meskalaensis nov. gen. nov. sp.
Besides the description of these new forms, the systematic assignment of many species is reevaluated.
Though the whole fauna is not studied in detail, all occuring taxa are considered for general questions. The exact range of different species allows to revise the stratigraphical scale established by Arambourg. Particularly, the Danian stage can be characterized by a typical faunal association. Maastrichtian and Ypresian can be divided into lower and upper parts on the basis of different faunal associations.
Dynamic and pattern of extinction and faunal revival of selachians are analyzed and permit us to identify two important events of high extinction rates: mass extinctions at the end of the Maastrichtian and selective extinctions (sharks) at the end of the early Ypresian.
Comparisons between moroccan faunas from contemporaneous ones from other regions of the world show continuous exchanges between these areas from Maastrichtian to Ypresian.
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